Rats have been shown to engage in multiple forms of reciprocal help and cooperation, including what is referred to as direct reciprocity—where a rat will help another rat that has previously helped them.
【结构分析】
主干部分:
Rats have been shown to engage in multiple forms of reciprocal help and cooperation
修饰成分:
including……为伴随状语;
what is referred to as direct reciprocity为动名词including后的宾语从句;
where a rat will help another rat为定语从句修饰direct reciprocity;
that has previously helped them为定语从句修饰another rat;
【译文】
有研究表明,老鼠会进行多种多样形式的互助及合作,其中包括被称为直接互惠的行为,即老鼠会帮助曾经帮助过自己的另一只老鼠。